Rory Putman addresses the question of how, in many temporate ecosystems, diverse and species-rich assemblies of ungulates manage to co-exist despite often quite extensive overlap in ecological requirements. Putman explores the potential for competition, competition tolerance and even positive facilitation amongst the members of such guilds of ungulates. As a central worked example, the author employs data resulting from over 20 years of personal research into the ecology and population dynamics of various large herbivores of the New Forest in Southern England. With these, he applies formal prools in resource use, evidence for resource limitation and evidence for interaction between species in changing population size over the years.